In Reply To:
In Reply To:
The contractile efficiency of the muscle is around 40%. The overall pedaling efficiency of the cyclist is about 20%. Account for all the losses without invoking losses from the pedaling motion itself.
I don't know, where these numbers come from and if they are valid. Let's assume, they are ok.
Actually, they aren't valid, for the following reasons:
1) the efficiency of glucose oxidation (which is where the ~40% value arises) is calculated based on standard conditions, which do not exist
in vivo. In particular, the actual delta G of ATP synthesis/hydrolysis is less than the assumed delta G zero prime, meaning that the actual efficiency of the initial energy 'capture' is <40%.
2) while the calculated efficiency of glucose oxidation may be ~40%, that is not the same as the efficiency of muscle contraction
per se. The latter is difficult to determine, but must be less than 100% (2nd law of thermodynamics), meaning that the overall efficiency must be <40%, even if the conversion of force at the myofibrillar level to useful external work occurs w/o any loss whatsoever.
3) although whole-body efficiency when cycling is around 20-25%, that is for the body as a whole, not for the exercising muscles themselves. (As a general rule-of-thumb, the percentage of whole-body VO2 consumed by the legs during cycling is approximately equal to the percentage of VO2max, e.g., at 70% of VO2max the legs account for ~70% of whole-body O2 uptake.) If you calculate efficiency based on leg instead of whole-body VO2, you get a significantly higher value, i.e., around 30-35%.
Putting 1-3 together, the picture that emerges is that humans are actually quite efficient when pedaling a bicycle, something that makes perfect sense when you realize that the pattern of muscle use/activation when pedaling is quite similar to that observed when walking or running, i.e., we use our muscles to pedal pretty much the way they were evolutionarily-designed to be used.