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In regards to the 405CX, it's an improved 405 with some extra features, I believe it is higher precision GPS and has improved HR monitoring capabilities. Overall, having owned both a 405 and now the 310xt, I really have a hard time recommending the 405 to any multisport athletes, I find it's really just too limited in terms of the interface, display, and overall applicability to multisport. Hence, I got rid of mine.
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405CX is basically the same thing as the 405. Two benefits- one is a velcro band for people with smaller wrists. The second is a set of European charging cables. I don't think the actual guts of the watch are any different. Oh, you do also get a fancy blue watchband. So three benefits!
The 310 won't transmit HR while you are actually underwater. The water prevents transmission. However, they will sync up afterwards and you can see your HR after the workout.
From the owners manual, page iv:
So, if you're a relatively fast swimmer or don't mind stretching the specs just a bit, it should be fine for a HIM swim.
Note that in my experience the bezel absolutely freaks out when its wet. Even rain or sweat can do it in. And if you have it set to backlight on key action (including bezel touches), this can drain the battery in about 15 minutes. So I'd definitely lock the bezel before the swim. If you want more than one screen of data, turn on auto scroll. That way you won't need to touch the bezel for anything.
Also note that the 405 doesn't have a multisport mode, so you'll have to go through a few button presses (including unlocking and then relocking the bezel) in transition.
From what I've heard the garmin units rely on GPS solely for altitude which has a lot of variance.
The bezel issues seem like a real PIA with the Garmins as well.
IPX7 is designed to withstand accidental immersion for that period of time. It is not designed for regular use in water. Certainly not for swimming Even if it does not leak the first time you try this (and it has for some people) it eventually will leak if you continue to use it in water. Garmin themselves make this clear. It should not be fine at all for your HIM swim unless you are lucky.
Please do not risk your expensive watch for something it was not designed to do. If you want waterproof Garmin, either buy the 310 or wait for the upcoming waterproof version of the 405 that is supposedly coming out.